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Giant tree-rat
Conservation status
Scientific classification
Genus:
Toromys
Species:
grandis
Synonyms

Loncheres grandis Wagner, 1845
Echimys grandis (Wagner, 1845)
Makalata grandis (Wagner, 1845)

The giant tree-rat (Toromys grandis) is a species in the family Echimyidae, the spiny rats. It is the only species in the monotypic genus Toromys. It is endemic to Brazil, where it occurs in the flooded forest along the banks of the Amazon River and its tributaries.

This species was formerly considered to be a member of the genera Loncheres and Makalata. In 2005 it was reclassified as the only member of the new genus Toromys.

The etymology of the genus name derives from the onomatopoeic Portuguese word toró, referring to the vocalizations of this rodent, and the ancient greek word μῦς (mûs), meaning "mouse, rat".

Phylogeny

Toromys is the sister genus to Pattonomys. These taxa are closely related to Echimys, Phyllomys, and Makalata, reflecting the fact that Toromys grandis and Pattonomys occasius have formerly been placed in Makalata by some authorities. In turn, these five genera share phylogenetic affinities with a clade containing the bamboo rats Dactylomys, Olallamys, Kannabateomys together with Diplomys and Santamartamys.

Genus-level cladogram of the Echimyini.
root  
         
         
         
         
         

  Echimys



  Phyllomys




  Makalata




         

  Pattonomys



  Toromys
  (Giant tree-rat)





  "Dactylomyines"  
         

  Dactylomys



  Olallamys




  Kannabateomys
  (Atlantic bamboo rat)





  Diplomys



  Santamartamys
  (Red-crested tree-rat)





  "Eumysopines"  

  Lonchothrix
  (Tuft-tailed spiny tree-rat)



  Mesomys





  Isothrix



The cladogram has been reconstructed from mitochondrial and nuclear DNA characters.
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